Washington State finished with 224 yards total offense.

Washington State came out passing as expected, with Tuel starting 8 of 9. But his 10th pass was intercepted by Uona Kaveinga, and the momentum shifted to BYU before a "white-out" crowd.

Andrew Furney's 47-yard field goal put Washington State on the board with 10:42 left in the second quarter and he added a 41-yarder just before halftime after a TD pass to Marquess Wilson was nullified by holding.

Washington State (1-1), which lost at BYU in the season opener, also unveiled a $65 million upgrade of Martin Stadium in front of the first sellout crowd here since 2007.

This was the first meeting since 1908 between the two programs, located 60 miles apart.

Washington State has never lost in 17 games against teams from the FCS.

Starting in place of the injured Jeff Tuel, Halliday threw two touchdown passes to Marquess Wilson, and also teamed with Gabe Marks and Isiah Myers on first-half scoring strikes.

Halliday completed 26 of 45 passes for 378 yards and had two interceptions.

Marks led the Cougars with six catches for 126 yards, and Wilson had five receptions for 110 yards.

Connor Halliday threw four touchdown passes for Washington State (2-2), which was looking for its first three-game winning streak since 2006.

Washington State was ahead 34-28 when Colorado took over on its 30 and marched to the WSU 4. On fourth down, Webb bootlegged up the middle for a touchdown to tie the score. Will Oliver's extra point gave Colorado its first lead, 35-34, with 9 seconds left.

The game was played in air thick with smoke from wildfires 200 miles away in central Washington.

Washington State continued its series of playing once a year at the home of the Seattle Seahawks and for the foreseeable future it will be either Oregon or Oregon State as the foe. While the crowd of 60,929 was mostly crimson, there was plenty of green and highlighter yellow that made the trip north to see the Ducks first road game of the season.

Connor Halliday threw for 348 yards and Marquess Wilson had 12 catches for 182 yards and a touchdown to become Washington State's all-time leader in yards receiving.

Jeff Tuel was 11 of 17 for 126 yards after replacing Connor Halliday, who threw three interceptions in just over one half of action. Marquess Wilson had four catches for 54 yards to lead the Cougars (2-4, 0-3). Halliday was 9-for-20 passing.

Turnovers kept the Cougars from gaining a foothold as they continue to implement new coach Mike Leach's pass-happy offense. Oregon State kept both Halliday and Tuel on the run, forcing four interceptions and a fumble. Defensive end Scott Crichton tallied three sacks.

C.J. Anderson rushed for two touchdowns as the Golden Bears (3-4, 2-2 Pac-12) earned their eighth straight win over the Cougars.

Washington State coach Mike Leach is still looking for his first Pac-12 win.

Jeff Tuel replaced Connor Halliday in the first half and threw for 320 yards and two touchdowns - both to Brett Bartolone.

WSU receiver Marquess Wilson, who caught four passes for 87 yards in the first half, was injured late in the half and did not return.

If not for its defense, Stanford's Pac-12 North Division hopes might be dashed.

Tuel led a 12-play, 75-yard drive capped by a 10-yard touchdown pass to Kristoff Williams with 6:29 to play to bring Washington State within a touchdown. Stanford managed two first downs before punting the ball back to Washington State.

The loss was the sixth straight for Washington State (2-7, 0-6 Pac-12), assuring coach Mike Leach of his first losing season.

Washington State's Jeff Tuel was 23 of 45 for 232 yards, with a 5-yard TD pass to Kristoff Williams on the game's final play. Tuel was intercepted, fumbled and sacked six times.

Tuel, who set a school passing mark with 43 completions last week as Washington State took it down to the wire against nationally ranked Stanford, completed three of his first five passes for 41 yards Saturday, but was just 1 of 9 for 6 yards the rest of the quarter.

UCLA (8-2, 5-2 Pac-12) has won four games in a row since a loss at California, and is first in the Pac-12 South.

Connor Halliday threw five touchdown passes for Washington State (2-8, 0-7) in the loss.

Washington State dominated the first quarter, then was outscored 30-0 in the second quarter as Hundley threw three touchdown passes and the Bruins piled up 181 yards.

Arizona State (6-5, 4-4 Pac-12) had failed in four tries to become bowl eligible, bogging down against some of the best teams in the conference.

The Sun Devils finally broke through behind their quarterbacks and a resurgent defense.

The Pac-12's passing leader, Washington State had 240 yards passing - 137 below its season average.

Late in the fourth quarter, Connor Halliday found Kristoff Williams for a 54-yard touchdown for the Cougars' only score.

Deone Bucannon matched a career-high with 17 tackles.

Washington quarterback Keith Price threw an interception on the first play of overtime that defensive lineman Kalafitoni Pole nearly returned for a game-ending touchdown, but he was caught at the 5.

The Cougars (3-9, 1-8 Pac-12) took possession and, after picking up a first down, Furney kicked a 27-yard field goal, leading to a crush of crimson and gray fans storming and celebrating at midfield.

In a rivalry known for unpredictability, the 2012 version will rank among the tops in Apple Cup lore.